
Research : Since 1971: Museum and laboratory research on the functional morphology, development and evolution of Malagasy lemurs; allometry and heterochrony. Since 1986: Field expeditions to Madagascar (Anjohibe, Anjohikely, Antsiroandoha, Andrafiabe, Fitsangatsanganan Ilay Olon' Tokana, Ampasambazimba, Taolambiby, Andrahomana and other sites), to explore the giant extinct lemur and associated fauna. Since 2001: Beza Mahafaly Special Reserve sifaka and mouse lemur projects. Ranomafana National Park lemur ontogeny project.