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Lloyd Gamble

Graduate Student (PhD)


Lloyd Gamble earned his PhD from the Department of Environmental Conservation at the University of Massachusetts in 2007, where he studied metapopulation dynamics, dispersal and viability in pond-breeding amphibians. He received his BA from the Department of Environmental Conservation at Vassar College in 1989. He is currently working with the World Wildlife Fund in Washington, DC.

Lloyd's MS and PhD research in the Landscape Ecology Lab focused on metapopulation dynamics of the marbled salamander (Ambystoma opacum) in western Massachusetts. Llyod was was instrumental in establishing the foundation for this long-term study. [Click here to learn more about the vernal pool amphibian project]

Lloyd's research focused on the following objectives: 1) characterizing the range of demographic variation and temporal synchrony among local marbled salamander populations, and 2) quantifying dispersal rates among breeding sites and subsequently, the degree of effective isolation or interaction among these sub-populations.

Lloyd's PhD work resulted in the following publications:

  • Gamble, L.R., K. McGarigal, D.B. Sigourney, and B.C. Timm. 2009. Survival and breeding frequency in Marbled Salamanders (Ambystoma opacum): implications for spatio-temporal population dynamics. Copeia 2:394-407. (pdf)
  • Gamble, L. R., S. Ravela, and K. McGarigal. 2008. Multi-scale features for identifying individuals in large biological databases: an application of pattern recognition technology to the marbled salamander Ambystoma opacum. Journal of Applied Ecology 45:170–180. (pdf)
  • Gamble, L.R., K. McGarigal, and B.W. Compton. 2007. Fidelity and dispersal in the pond-breeding amphibian, Ambystoma opacum: Implications for spatio-temporal population dynamics and conservation. Biological Conservation 139:247-257. (pdf)
  • Gamble, L. R., K. McGarigal, C. L. Jenkins, and B. C. Timm. 2006. Limitations of regulated “buffer zones” for the conservation of marbled salamanders. Wetlands 26(2):298-306. (pdf)

For more information, please contact:
Dr. Kevin McGarigal
Department of Environmental Conservation
University of Massachusetts
304 Holdsworth Natural Resources Center
Box 34210, Amherst, MA 01003
Fax: (413) 545-4358; Phone: (413) 577-0655

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